The Mark V was first to be a completely new design. When however in December 1917 the desired new engine and transmission became available, this design was abandoned and the designation switched to an improved version of the Mark IV, in fact a Mark IV as it was originally intended: more power (150 bhp) with a new Ricardo engine, improved steering mechanism and epicyclical transmission, only one driver was needed. Cabin for machine-gunner on the roof. 400 were built, 200 each of Males and Females. Several were converted to Hermaphrodites by swapping sponsons to give a single 6 pounder gun for each.